Bothremydidae

Bothremydidae Temporal range: 99.6–40.4 Ma PreЄ Є O S D C P T J K Pg N
Gafsachelys spp. skull
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Pleurodira
Family:	†Bothremydidae Baur, 1891

Bothremydidae is an extinct family of side-necked turtles. They belong to suborder Pleurodira and the order Testudines. Like their modern relatives, bothremydidids often inhabited freshwater, but several species were instead marine;^[1] they are unrelated to modern sea turtles. They were actively mobile omnivores. They became extinct in the Late Eocene.^[2]

The family is split into two subfamilies and a number of tribes.^[3]

Bothremydidae

• Subfamily Bothremydinae Gaffney et al. 2006
  • Tribe Bothremydini Gaffney et al. 2006
    • Subtribe Bothremydina Gaffney et al. 2006
      • Araiochelys Gaffney et al. 2006
        • Araiochelys hirayamai Gaffney et al. 2006
      • Bothremys Leidy 1865 (jr synonym: Karkaemys)
        • Bothremys arabicus (Zalmout et al. 2005)
        • Bothremys cooki Leidy 1865
        • Bothremys kellyi Gaffney et al. 2006
        • Bothremys maghrebiana Gaffney et al. 2006
      • Chedighaii Gaffney et al. 2006
        • Chedighaii barberi (Schmidt 1940)
        • Chedighaii hutchisoni Gaffney et al. 2006
      • Rosasia Costa 1940
        • Rosasia soutoi Carrington da Costa 1940
      • Inaechelys Carvalho 2016
        • Inaechelys pernambucensis Caralho 2016
      • Zolhafah Lapparent de Broin and Werner 1998
        • Zolhafah bella Lapparent de Broin and Werner 1998
    • Subtribe Foxemydina Gaffney et al. 2006
      • Elochelys Nopsca 1931
        • Elochelys convenarum Laurent et al. 2002
        • Elochelys perfecta Nopsca 1931
      • Foxemys Tong et al. 1998
        • Foxemys mechinorum Tong et al. 1998
      • Polysternon Portis 1882
        • Polysternon atlanticum Lapparent de Broin and Murelaga 1996
        • Polysternon provinciale (Matheron 1869)
      • Puentemys Cadena et al., 2012
        • Puentemys mushaisaensis Cadena et al., 2012
  • Tribe Taphrosphyini Gaffney et al. 2006
    • Subtribe Nigermydina Gaffney et al. 2006
      • Arenila Lapparent de Broin and Werner 1998
        • Arenila krebsi Lapparent de Broin and Werner 1998
    • Subtribe Taphrosphyina Gaffney et al. 2006
      • Azabbaremys Gaffney et al. 2001
        • Azabbaremys moragjonesi Gaffney et al. 2001
      • Phosphatochelys Gaffney and Tong 2003
        • Phosphatochelys tedfordi Gaffney and Tong 2003
      • Rhothonemys Gaffney et al. 2006
        • Rhothonemys brinkmani Gaffney et al. 2006
      • Taphrosphys Cope 1869 (jr synonyms: Amblypeza, Bantuchelys, Prochonias)
        • Taphrosphys congolensis (Dollo 1912)
        • Taphrosphys dares Hay 1908 (nomen dubium)
        • Taphrosphys ippolitoi Gaffney et al. 2006
        • Taphrosphys sulcatus (Leidy 1856)
      • Ummulisani Gaffney et al. 2006
        • Ummulisani rutgersensis Gaffney et al. 2006
      • Crassachelys (nomen dubium)
        • Crassachelys neurirregularis (Bergounioux 1952)
      • Eusarkia (nomen dubium)
      • Gafsachelys Stefano 1903 (nomen dubium)
      • Labrostochelys Gaffney et al. 2006
        • Labrostochelys galkini Gaffney et al. 2006
  • Tribe Cearachelyini Gaffney et al. 2006
    • Cearachelys Gaffney et al.
      • Cearachelys placidoi Gaffney et al. 2001
    • Galianemys Gaffney et al. 2002
      • Galianemys emringeri Gaffney et al. 2002
      • Galianemys whitei Gaffney et al. 2002
• Subfamily Kurmademydinae Gaffney et al. 2006
  • Tribe Kurmademydini Gaffney et al. 2006
    • Kinkonychelys Gaffney et al. 2009
      • Kinkonychelys rogersi Gaffney et al. 2009
    • Kurmademys Gaffney et al. 2001
      • Kurmademys kallamedensis Gaffney et al. 2001
    • Sankuchemys Gaffney et al. 2003
      • Sankuchemys sethnai Gaffney et al. 2003

Phylogeny

Below is a cladogram by Gaffney et al. in 2006:^[4]

References